Typically a woodland butterfly, found on or near oaks (its larval food plant), it may also be seen in parkland, large gardens, or hedgerows. It favours trees with a south-facing aspect, along rides, woodland edges and clearings where it is sheltered and sunny. It can be found on either of our native oaks, as well as on introduced species.
An elusive butterfly, it spends much of its time in the treetops where it is best observed through binoculars although it will occasionally descend to ground level. It is often active on warm, sunny evenings, when large numbers may be seen at favoured sites, although it has been noted that if the sun disappears behind clouds all activity suddenly ceases.
Like the White-letter Hairstreak this species feeds on honeydew secreted by aphids, which collects on the surface of leaves, only rarely nectaring on flowers.
There is a single record of an individual seen on moorland in VC62 well away from any areas of woodland, suggesting that this species may be capable of dispersing a considerable distance.